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- Traits:Biodiffusor + (Organisms whose activities that cause cons … Organisms whose activities that cause constant and random local sediment biomixing over short distances resulting in transport of sediment particles, analogous to molecular or eddy diffusion (from Kristensen ''et al.'', 2012). Includes epifaunal biodiffusers e.g. fiddler crabs; surficial biodiffusers e.g. ''Echinocardium''; and gallery biodiffusers e.g. ''Nereis (Hediste) diversicolor''.rs e.g. ''Nereis (Hediste) diversicolor''.)